The game will be played at the DW Stadium in Wigan, Tomkins’ home ground with the Warriors.
Huddersfield prop Eorl Crabtree has been named on the bench by Tony Smith.
Tomkins is partnered at half-back by Grand Final winning stand-off Danny McGuire, who is one of five Leeds players in the starting line-up – together with Lee Smith, Ryan Hall, Jamie Peacock and Kevin Sinfield.
Meanwhile, Australia coach Tim Sheens has called up experienced back-rower Nathan Hindmarsh for the clash, and will also hand debuts to hooker Robbie Farah, hooker Luke Lewis and prop David Shillington.
England: S Briscoe (Hull KR); T Briscoe (Hull), Smith (Leeds), Shenton (Castleford), Hall (Leeds); McGuire (Leeds), Tomkins (Wigan); Morley (Warrington), Roby (St Helens), Graham (St Helens), Ellis (Wests Tigers), Peacock (Leeds, captain), Sinfield (Leeds). Substitutes: Burgess (Bradford), Crabtree (Huddersfield), Eastmond (St Helens), Westwood (Warrington).
Australia: Slater; B Morris, Inglis, Hodges, Hayne; Lockyer (capt), Thurston; Civoniceva, Smith, Hannant, Watmough, Gallen, Hindmarsh. Replacements (from): Farah, White, Lewis, Shillington, Waterhouse, Gidley.